The film then moves on to Nemo's first day of school. Nemo has a small right fin (due to a minor injury to his egg from the barracuda attack) which causes Marlin to not only worry over his swimming ability but also doubt his son’s general capabilities. After Marlin embarrasses Nemo during a school field trip, Nemo disobeys his father and sneaks away from the reef towards a boat, resulting in him being captured by a scuba diver. However, as the boat sails away, the diver accidentally knocks his diving mask into the water.
Bruce, a great white shark; Anchor, a hammerhead shark; and Chum, a mako shark, Marlin discovers the diver's mask that was dropped from the boat and notices an address written on it. However, he struggles with Dory and the mask snaps against Dory's nose, causing a nosebleed. The scent of blood causes Bruce to lose control of himself, and he attempts to eat Marlin and Dory. Shortly after the two escape from Bruce, the mask is dropped into a trench in the deep sea. During a hazardous struggle with an anglerfish in the trench, Dory realizes she is able to read the address written on the mask, which leads to Sydney, Australia, and manages to remember it despite her short-term memory loss. After receiving directions from a large school of moonfish, Marlin and Dory set out to find Sydney. After a struggle with jellyfish that nearly sting Dory and him to death, Marlin falls exhausted after the risky escape and wakes up to see a surf-cultured sea turtle named Crush, who takes Dory and him on the East Australian Current. Marlin sees Dory unconscious, thinks she is dead and blames himself, but she wakes up also alive and well. In the current, Marlin reluctantly shares the details of his journey with a group of young sea turtles, and his story spreads rapidly across the ocean through word of mouth and eventually reaches Nemo in Sydney.
Upon leaving the East Australian Current, Marlin and Dory become lost in a huge bloom of plankton and krill, and are engulfed by the mouth of a whale after swimming in its path. Inside the whale's immense mouth, a desperate Marlin repeatedly charges against the baleen in the whale's mouth in a futile attempt to free himself and Dory, then breaks down in despair. Dory reassures him, then calmly tries to communicate with the whale. In response, the whale carries them to Sydney Harbour and expels them through its blowhole. They are then met by Nigel, who recognizes Marlin from the stories he has heard and rescues him and Dory from a flock of hungry seagulls by scooping them into his beak and taking them to the dentist's office. By this time, Darla has arrived and the dentist is prepared to give Nemo to her. Nemo tries to play dead in hopes of saving himself, and, at the same time, Nigel arrives. Marlin sees Nemo and mistakes this act for the actual death of his son. After the dentist throws Nigel out, Gill helps Nemo escape into a drain through a sink plug-hole after a chaotic struggle.
Back at the dentist's office, the high-tech filter breaks down and The Tank Gang have escaped into the harbor, but realize they are confined to the bags of water that the dentist put them into when cleaning the tank.
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